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Recent advances in the chemistry and biochemistry of plant lipids, Academic Press, New York, pp. 247-286. 10. V. G. (1980) Fisiologiya rastenii, 27, 390-398. 11. P. G. (1987) Khimiya prir. 1, 52-57. 12. Litchfield, C. (1972) Analysis of triglycerides, Academic Press, New York, 355 pp. 36 BIOlOOICAI. A. aw:s, It OIMZ, T. KABIIoIEII (ocIo~ A k _ KIId6, BudIpooIIllld _ ~ Cofporotton, _ V.... IIId london, , .... J. Mwphy and I. Cwnmlna Department of BIologIcal Sciences, UnlversHy of Durham, DH1 3lE,UK INTRODUCTION Storage triacylglycerols are broken down by triacylglycerol lipases following seed germination.

3% of the sum N+NE. 4%. 9 tr. 8 atter flowering Fig. 1. Dynamics of the mesocarp (M) and seeds (S) fresh weight and the absolute content of their total lipids, TAGs and polar lipids during maturation -"-, -~- total and "defatted" fresh weight; - - , _ . - total and polar lipids;--- TAGs 33 s M z /'/' .. ~ fJ' ...... • ~ o ~ .. ,. ig o . >II 10 Days "fter flowering Fig. 2. Dynamics of the absolute content of inPividual esterified FAs in maturing fruits. FAs: 1-palmitic, 2-hexadecenoic, 3-hexadecadienoic, 4-stearic, 5-oleic, 6-linoleic, 7-linolenic; ---- total lipid FAs, --- FAs of triacylglycerols.

In members of Boraginacede and CaryophyZZacede 20:4w 6 was not found too, but 18:3w 6 and 18:4w 3 are present 110, 11 I. Due to the absence of acids with ~5-double bond, it seems possible that 20:4w 6 forma- 46 tion in members of these two families is limited by the lack of ~5-desaturase activity. Up to date, there are no strongly evidential data suggesting the presence of 20:4w 6 and 20:5w 3 in flowering plants, where 18:3w3 is the main component. There are no data of biosynthetic experiments, confirming the presence of both ~6_ and ~5-desaturases in some higher plant.

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